请分析一下这句话中kill在这里是什么成分?或者who引导的从句中谓语动词是什么?

问题描述:

请分析一下这句话中kill在这里是什么成分?或者who引导的从句中谓语动词是什么?
"we owe a lot to birds and beasts who eat insects but all of them put together kill only a fraction of the number destroyed by spiders."

we owea lot to birds and beasts(who eat insects)——这是宾语从句.
but后面是并列句.
all of them (put together后置定语,put是过去分词) kill(谓语)
only a fraction of the number destroyed by spiders.