We owe a lot to the birds and beasts who eat insects but all of them put together kill only a fraction of the number destroyed by spiders. all of them 指的是什么,是它们杀死的昆虫的所有,还是它们本身即鸟兽的所有

问题描述:

We owe a lot to the birds and beasts who eat insects but all of them put together kill only a fraction of the number destroyed by spiders. all of them 指的是什么,是它们杀死的昆虫的所有,还是它们本身即鸟兽的所有

all of them ==the insects that birds and beasts eat

birds and beasts
owe 这里应该是欠。
我们欠那些吃虫子的鸟兽很多
owe表示“欠(债)”与介词for连用时,后面只接物; 与介词to连用时,后面可接人,也可接物或事物

指鸟兽的所有.
这句话意思:我们要归功于许多吃掉昆虫的鸟兽,但是所有这些鸟兽加起来消灭的昆虫只占蜘蛛消灭昆虫量的一小部分.
注意all of them后面谓语是kill,显然kill的主语是鸟兽,