he laughs best who laughs last.是什么从句

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• 请问“He who laughe last laughs best ”的中文意思是什么?
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• he laughs best ( )laughs last.这里的先行词是he吗?为什么不跟关系词紧挨着?
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• 【英语】：麻烦大神说下列下句子关系代词做什么成分?①i'll never forget the days which[状语?] we spent together in the countryside.②this is the best book that[状语?] i have ever read③the road conditions there turned out to be very good,which[宾语?]was more than we could expect④AS[主语?]is known to us all,china is becoming stronger and stronger⑤he who[主语?] laughs last laughs best⑥the boy whom[宾语?] i am looking after is my brother's son⑦we must help those who[主语?] need help⑧i met a boy wh
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