What constitutes a major threat to the survival of turtles according to Elizabeth Griffin?

问题描述:

What constitutes a major threat to the survival of turtles according to Elizabeth Griffin?
此句的主语是what吗

What constitutes a major threat to the survival of turtles according to Elizabeth Griffin?根据伊丽莎白·格里芬,什么对海龟的继续生存构成了主要威胁?疑问词 what 就是这句话的主语,谓语动词是 constitute,其...